Okay, Here's the Explanation
I'm an idiot.When you play bonus songs or downloadable content songs, they get added to your offline campaign score. I didn't know that, because I hadn't played any bonus songs in the 360 version yet.
I played two DLC songs in the last day or so, and when it added those scores to my campaign score, I didn't even notice. Later, when I checked the leaderboards, my offline and online scores didn't match.
I verified that the DLC songs I've played account for the score discrepancy to the point. So that's clearly what happened.
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